Here are 22 Photo Masterclasses written by Mark to provide detailed practical advice on many fields of wildlife and nature photography, from mammal portraits and bird behaviour to black and white and different photographic techniques (click on each below). Scroll to the bottom for articles about how to win the Wildlife Photographer of the Year competition, camera advice and the ethics of digitally manipulated wildlife photography (all originally published in BBC Wildlife magazine).

Mammal Portraits	Urban & Garden	Invertebrates	Extreme Close-up	Animals in their Environment	Creative Visions	Cold Weather Wildlife
Wet Weather	Underwater	Mammal Behaviour	Bird Behaviour	Dusk to Dawn	Wild Places	Black & White
Plant Portraits	Zoo Photography	Reptiles & Amphibians	The World in our Hands	Bird Portraits	Birds in Flight	Innovative Wildlife Photography
Keep it Simple	Which SLR camera?	Which compact camera?	How to win Wildlife Photographer of the Year	Wildlife Photographer of the Year - an insider's guide	The ethics of digital wildlife photography